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My reference is a pair of Beyerdynamic DT 880 SE (600 ohms). The HE4XX has louder and more defined mids (vocal), and more distinct bass percussions. The sound stage seems smaller and flatter. So far, I feel it plays nicer with vocal music. For the price, I can't complain.
I am driving it with Schiit Vali 2 on low gain with two different tubes (Tung-Sol 6SN7GTB and Genalex E88CC Gold Lion), but in a pinch it is possible to drive with a cellphone or iPad.
I have now also tried the HE 4XX with Schiit Fulla 2. Compare to Vali 2, it sounded slightly "coarser", but we are talking about a very small difference.